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# of Endemic Species
809: Bass Strait Drainages
Major Habitat Type:
temperate coastal rivers
This ecoregion covers the northern half of Tasmania from the Ansons River west to the Welcome River, and a portion of the coast on mainland Australia that extends from Wilson’s Promontory west to the edge of the Murray-Darling Basin.
Drainages flowing into:
Main rivers or other water bodies:
Several coastal rivers including the Tamar, Mersey, Forth, (all in Tasmania), Tarwin, Bunyip, Yarra, Barwon, Lake Corangamite, Hopkins and Glenelg occur within this ecoregion.
On the mainland, elevations reach a maximum of about 600 m in the hills at the northern edge of the ecoregion. Elevations are higher, up to a maximum of about 1300 m, in the highlands of Tasmania. In both regions the elevations drop off quickly moving towards the coast.
Rainfall is relatively high in this part of Australia with the highest precipitation on Tasmania.
This ecoregion is relatively small, covering 105,000 square kilometers or 1.3% of Australia. From the Yarra River east, coastal rivers descend from higher forested catchments and tend to have perennial flow. Rivers west of the Yarra rise in lower elevation headwaters with low gradients and portions of these drainages are often intermittent during dry periods. Most of the Tasmanian rivers are short and flow north from into Bass Straight. This is one of only two Australian ecoregions that has a significant number of permanent natural lakes. These are mostly situated in the western part of the mainland portion of the ecoregion. Some are situated in volcanic craters and tend to be deep, while others are much shallower and tend to be slightly saline.
A diverse range of vegetation types occurs in this ecoregion, including coastal vegetation, temperate rainforest, and open eucalyptus woodlands (World Wildlife Fund 2001a, 2001b).
The fish fauna of this ecoregion is dominated by the family Galaxiidae (Galaxiids), followed by Percichthyidae (temperate perches). The remaining six families have three species or fewer.
Description of endemic fishes:
Three species are endemic to this province: Galaxias tanycephalus (saddled galaxias), Paragalaxias mesotes (Arthurs paragalaxias), and Nannoperca variegata (variegated pygmy perch). Galaxias tanycephalus is a land-locked derivative of the diadromous G. truttaceus, spotted galaxias (Waters et al. 2000).
Other noteworthy fishes:
While precise research is lacking, it appears likely that Galaxiella pusilla, dwarf galaxias, may be capable of aestivation (Beck 1985). Neochanna cleaveri, the Australian mudfish, is another galaxiid fish that is also capable of aestivation (McDowall & Fulton 1996). Both species are found also in neighboring ecoregions.
Other noteworthy aquatic biotic elements:
The crayfish genus Euastacus is restricted to the Australian mainland, and five species are found in the northern (mainland) section of the ecoregion (Morgan 1986). Two of these species (E. bispinosus and E. woiwuru) are endemic to the ecoregion. The giant freshwater lobster, Astacopsis gouldi, is endemic to the Tasmanian section of the ecoregion (Hamr 1992; Horwitz 1994). The monotypic genus Gramastacus is restricted to a small area of the Grampians region (Zeidler & Adams 1990). The genus Geocharax is currently endemic to the ecoregion, occurring on both the Tasmanian and mainland sections, although an undescribed species is also known from Eastern Coastal Australia  (Horwitz 1995a). Cherax destructor occurs naturally in the mainland section of the ecoregion (Sokol 1988) and has also been translocated into the Tasmanian section (Elvey et al. 1996). Twenty-nine species of the crayfish genus Engaeus occur in the ecoregion. Seven of these species (E. australis, E. fultoni, E. sericatus, E. sternalis, E. strictifrons, E. urostrictus and E. victoriensis) are endemic to the mainland portion, and nine (E. granulatus, E. leptorhynchus, E. mairener, E. martigener, E. nulloporius, E. orramakunna, E. spinicaudatus, E. tayatea and E. yabbimunna) are endemic to the Tasmanian portion (Horwitz 1990, 1995b).
Justification for delineation:
Although the mainland and northern Tasmania are currently separated by the Bass Straight, they were connected during lowered sea-levels and thus share a similar fauna. The ecoregion is distinctive relative to Eastern Coastal Australia  in the species that it lacks, likely due to a former drainage divide between these regions during lowered sea levels. Despite a lack of significant topographic barriers, the fauna of the Murray-Darling ecoregion contains a number of species lacking from this ecoregion as well (Unmack 2001).
Level of taxonomic exploration:
Overall taxonomic exploration is good. However, many species within this province remain poorly studied and recent evidence suggests several undescribed taxa exist within more widespread forms. At least four fish species may require revisions that may increase the number of species (e.g., Galaxiella pusilla (Unmack & Adams unpub. data), Galaxias olidus (Raadik unpub. data), Retropinna semoni (Hammer et al. 2007) and Gadopsis marmoratus (Miller et al. 2004)).
Beck, R. G. (1985). "Field observations upon the dwarf galaxiid Galaxiella pusilla (Mack) (Pisces: Galaxiidae) in the south-east of South Australia" South Australia Naturalist 60 12-23.
Elvey, W., Richardson, A. M. M., et al. (1996). "Interactions between the introduced yabby, Cherax destructor, and the endemic crayfish Astacopsis franklinii, in Tasmanian streams" Freshwater Crayfish 11 349-363.
Hammer, M., Adams, M., et al. (2007). "A rethink on Retropinna: conservation implications of new taxa and significant genetic sub-structure in Australian smelts (Pisces: Retropinnidae)" Marine and Freshwater Research 58 327-341.
Hamr, P. (1992). "A revision of the Tasmanian freshwater crayfish genus Astacopsis Huxley (Decapoda: Parastacidae)" Papers and Proceedings of the Royal Society of Tasmania 126 91-94.
Horwitz, P. (1995)"Preliminary Key to the Species of Decapoda (Crustacea, Malacostraca) Found in Australian Inland Waters" In Identification Guide no. 5. Albury NSW: Murray Darling Freshwater Research Centre, Cooperative Research Centre for Freshwater Ecology.
Horwitz, P. (1990). "A taxonomic revision of species in the freshwater crayfish genus Engaeus Erichson (Decapoda: Parastacidae)" Invertebrate Taxonomy 4 427-614.
Horwitz, P. (1995). "The conservation status of Australian freshwater crayfish: review and update" Freshwater Crayfish 10 70-80.
Horwitz, P. (1994). "Distribution and conservation status of the Tasmanian giant freshwater lobster Astacopsis gouldi (Decapoda: Parastacidae)" Biological Conservation 69(199-206)
Horwitz, P.,Adams, M. (2000). "The systematics, biogeography and conservation status of the species in the freshwater crayfish genus Engaewa Riek (Decapoda: Parastacidae) from south-western Australia" Invertebrate Taxonomy 14 655-80.
McDowall,Fulton (1996)"Family Galaxiidae" In McDowall, R.M. (Ed.). Fishes of South-eastern Australia. (pp. 52-77) Chatswood, Australia: Reed Books.
Miller, A. D., Waggy, G., et al. (2004). "Mitochondrial 12S rRNA sequences support the existence of a third species of freshwater blackfish (Percichthyidae: Gadopsis) from south-eastern Australia" Memoirs of Museum of Victoria 61 121-127.
Sokol, A. (1988). "Morphological variation in relation to the taxonomy of the destructor group of the genus Cherax" Invertebrate Taxonomy 2 55-79.
Unmack, P. J. (2001). "Biogeography of Australian freshwater fishes" Journal of Biogeography 28(9) 1053-1089.
Waters, J. M., López, J. A., et al. (2000). "Molecular phylogenetics and biogeography of galaxiid fishes (Osteichthyes: Galaxiidae): dispersal, vicariance, and the position of Lepidogalaxias salamandroides" Systematic Biology 49 777-795.
World Wildlife, Fund (2001). "Southeast Australia temperate forests (AA0409)" 2005 (2005; www.worldwildlife.org/wildworld/profiles/terrestrial/aa/aa0409_full.html).
World Wildlife, Fund (2001). "Tasmanian temperate forests (AA0412)" 2005 (2005; www.worldwildlife.org/wildworld/profiles/terrestrial/aa/aa0412_full.html).
Zeidler, W.,Adams, M. (1990). "Revision of the Australian crustacean genus of freshwater crayfish Gramastacus Riek (Decapoda: Parastacidae)" Invertebrate Taxonomy 3 913-924.